Eukaryotic nuclear genomes are generally considered to be large and gene-sparse, but extreme reduction has taken place several times, resulting in small genomes with a high gene-density. This process involves losing genes, compacting those that remain, or often both. Recently sequenced nuclear genomes include several that have converged to similar gene-densities by many means: variation in numbers and lengths of genes, intergenic regions and introns all contribute, but not equally in any given…
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